Princess Mononoke’s mother wolf Moro and Howl’s Moving Castle’s Witch of the Waste are both voiced by a famous singer and drag queen Akihiro Miwa.
Akihiro Miwa (born in 1935) is one of the most prominent queer icon in Japanese history, who survived the atomic bombing of Nagasaki (his hometown) during WW2.
This clip is from a document How Princess Mononoke Was Born (1998)